Carolan, James C., Murray, Tomas E., Fitzpatrick, Una, Crossley, John, Schmidt, Hans, Cederberg, Bjorn, McNally, Luke, Paxton, Robert J., Williams, Paul H. and Brown, Mark J.F. (2012) Colour Patterns Do Not Diagnose Species: Quantitative Evaluation of a DNA Barcoded Cryptic Bumblebee Complex. PLoS ONE, 7 (1). e29251. ISSN 1932-6203
Williams, Paul H., Brown, Mark J.F., Carolan, James C., An, Jiandong, Goulson, Dave, Aytekin, A. Murat, Best, Lincoln R., Byvaltsev, Alexandr M., Cederberg, Bjorn, Dawson, Robert, Huang, Jiaxing, Ito, Masao, Monfared, Alireza, Raina, Rifat H., Schmid-Hempel, Paul, Sheffield, Cory S., Šima, Peter and Xie, Zenghua (2012) Unveiling cryptic species of the bumblebee subgenus Bombus s. str. worldwide with COI barcodes (Hymenoptera: Apidae). Systematics and Biodiversity, 10 (1). pp. 21-56. ISSN 1477-2000
Williams, Paul H., Brown, Mark J.F., Carolan, James C., An, Jiandong, Goulson, Dave, Aytekin, A. Murat, Best, Lincoln R., Byvaltsev, Alexandr M., Cederberg, Bjorn, Dawson, Robert, Huang, Jiaxing, Ito, Masao, Monfared, Alireza, Raina, Rifat H., Schmid-Hempel, Paul, Sheffield, Cory S., Šima, Peter and Xie, Zenghua (2012) Unveiling cryptic species of the bumblebee subgenus Bombus s. str. worldwide with COI barcodes (Hymenoptera: Apidae). Systematics and Biodiversity, 10 (1). pp. 21-56. ISSN 1477-2000