Scriven, Jessica J., Woodall, Lucy C., Tinsley, Matthew C., Knight, Mairi E., Williams, Paul H., Carolan, James C., Brown, Mark J.F. and Goulson, Dave (2015) Revealing the hidden niches of cryptic bumblebees in Great Britain: Implications for conservation. Biological Conservation, 182. pp. 126-133. ISSN 0006-3207
Williams, Paul H., An, Jiandong, Brown, Mark J.F., Carolan, James C., Goulson, Dave, Huang, Jiaxing and Ito, Masao (2012) Cryptic Bumblebee Species: Consequences for Conservation and the Trade in Greenhouse Pollinators. PLoS ONE, 7 (3). e32992. ISSN 1932-6203
Williams, Paul H., Brown, Mark J.F., Carolan, James C., An, Jiandong, Goulson, Dave, Aytekin, A. Murat, Best, Lincoln R., Byvaltsev, Alexandr M., Cederberg, Bjorn, Dawson, Robert, Huang, Jiaxing, Ito, Masao, Monfared, Alireza, Raina, Rifat H., Schmid-Hempel, Paul, Sheffield, Cory S., Šima, Peter and Xie, Zenghua (2012) Unveiling cryptic species of the bumblebee subgenus Bombus s. str. worldwide with COI barcodes (Hymenoptera: Apidae). Systematics and Biodiversity, 10 (1). pp. 21-56. ISSN 1477-2000
Williams, Paul H., Brown, Mark J.F., Carolan, James C., An, Jiandong, Goulson, Dave, Aytekin, A. Murat, Best, Lincoln R., Byvaltsev, Alexandr M., Cederberg, Bjorn, Dawson, Robert, Huang, Jiaxing, Ito, Masao, Monfared, Alireza, Raina, Rifat H., Schmid-Hempel, Paul, Sheffield, Cory S., Šima, Peter and Xie, Zenghua (2012) Unveiling cryptic species of the bumblebee subgenus Bombus s. str. worldwide with COI barcodes (Hymenoptera: Apidae). Systematics and Biodiversity, 10 (1). pp. 21-56. ISSN 1477-2000